Between September and November 2012, non-invasive surveys were conducted by dividing the national park into survey blocks based on watershed boundaries. Search transects followed mainly along trails in each block to collect and store fresh bear scats in 95% ethanol as described in Bellemain et al. 2007. Spatial locations of all scat samples were recorded by a handheld GPS. Camera trapping was conducted simultaneously at 116 stations along the same trails. The procedure is described in details in Bischof et al. 2014.